With city approval of a proposed mosque and Islamic center two blocks away from Ground Zero nearly secured, the battle over the project is moving away from zoning boards to the court of public opinion.

The expected clearing of another major regulatory hurdle on Tuesday is unlikely to silence the debate around an issue that has become a call to arms for national Republicans—who have been emboldened by polls showing widespread public skepticism about the project—and a divisive policy matter for religious groups.

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